In 19th-century Paris, young vampire Noé finds the "Book of Vanitas" in the hands of a doctor, who tempts Noé with a plan to cure all vampires.

Vanitas and Noé flee with the Chasseurs in hot pursuit; Vanitas reveals why he entered the Chasseur's underground domain and how he knows his way around; after an unnecessary amount of drama, Vanitas finds whom he seeks.

Vanitas reunites with Dr. Moreau, who speaks of his work and a powerful backer; Vanitas has a flashback to his time with the Vampire of the Blue Moon; Vanitas and Noé try to save a curse-bearer with Roland's help.

Dominique offers Jeanne advice; Jeanne surprises Vanitas with an invitation, and Dominique shadows them; Noé encounters Lord Ruthven, but he doesn't know what to expect; Jeanne keeps her promise, and Vanitas makes one to her.

Jeanne says something that strikes fear into Vanitas; Lord Ruthven makes Noé swear an oath; Olivier is at his wits' end over Roland's obsession; Vanitas hears a certain monster has returned and sets out with Noé to investigate.